diff --git a/CHANGELOG.md b/CHANGELOG.md
index bb383a79..946b3545 100644
--- a/CHANGELOG.md
+++ b/CHANGELOG.md
@@ -38,6 +38,7 @@ Any violations of this scheme are considered to be bugs.
 * [#516](https://github.com/hashie/hashie/issues/516): Fixed `NoMethodError` raised when including `Hashie::Extensions::Mash::SymbolizeKeys` and `Hashie::Extensions::SymbolizeKeys` in mashes/hashes with non string or symbol keys - [@carolineartz](https://github.com/carolineartz).
 * [#531](https://github.com/hashie/hashie/pull/531): Fixed [slice doesn't work using symbols](https://github.com/hashie/hashie/issues/529) using hash with `IndifferentAccess` extension - [@gnomex](https://github.com/gnomex).
 * [#533](https://github.com/hashie/hashie/pull/533): Fixed `NoMethodError: undefined method `to_json'` at `hashie/dash_spec` - [@gnomex](https://github.com/gnomex).
+* [#535](https://github.com/hashie/hashie/pull/535): Restored the exporting of all properties as part of `Dash#to_h` and `Dash#to_hash` - [@michaelherold](https://github.com/michaelherold).
 * Your contribution here.
 
 ### Security
diff --git a/lib/hashie/dash.rb b/lib/hashie/dash.rb
index 460790b2..89a77a15 100644
--- a/lib/hashie/dash.rb
+++ b/lib/hashie/dash.rb
@@ -169,6 +169,13 @@ def replace(other_hash)
       self
     end
 
+    def to_h
+      defaults = ::Hash[self.class.properties.map { |prop| [prop, self.class.defaults[prop]] }]
+
+      defaults.merge(self)
+    end
+    alias to_hash to_h
+
     def update_attributes!(attributes)
       update_attributes(attributes)
 
diff --git a/lib/hashie/extensions/dash/indifferent_access.rb b/lib/hashie/extensions/dash/indifferent_access.rb
index 204cae6d..50d4f935 100644
--- a/lib/hashie/extensions/dash/indifferent_access.rb
+++ b/lib/hashie/extensions/dash/indifferent_access.rb
@@ -19,6 +19,15 @@ def self.requires_class_methods?(klass)
         end
         private_class_method :requires_class_methods?
 
+        def to_h
+          defaults = ::Hash[self.class.properties.map do |prop|
+            [Hashie::Extensions::IndifferentAccess.convert_key(prop), self.class.defaults[prop]]
+          end]
+
+          defaults.merge(self)
+        end
+        alias to_hash to_h
+
         module ClassMethods
           # Check to see if the specified property has already been
           # defined.
diff --git a/spec/hashie/dash_spec.rb b/spec/hashie/dash_spec.rb
index 387f0cb1..20ffb331 100644
--- a/spec/hashie/dash_spec.rb
+++ b/spec/hashie/dash_spec.rb
@@ -416,6 +416,7 @@ class KeepingMash < Hashie::Mash
         Class.new(Hashie::Dash) do
           property :a, required: -> { b.nil? }, message: 'is required if b is not set.'
           property :b, required: -> { a.nil? }, message: 'is required if a is not set.'
+          property :c, default: -> { 'c' }
         end
       end
 
@@ -434,6 +435,40 @@ class KeepingMash < Hashie::Mash
       it 'raises an error when neither property is set' do
         expect { codependent.new(a: nil, b: nil) }.to raise_error(ArgumentError)
       end
+
+      context 'exporting nil values' do
+        describe '#to_h' do
+          it 'does not prune nil values' do
+            expect(codependent.new(a: 'hi', b: nil).to_h).to eq(a: 'hi', b: nil, c: 'c')
+            expect(codependent.new(a: 'hi', b: nil, c: nil).to_hash).to eq(a: 'hi', b: nil, c: 'c')
+            expect(codependent.new(a: 'hi', b: nil).merge(c: nil).to_h).to eq(a: 'hi', b: nil, c: nil)
+          end
+        end
+
+        describe '#to_hash' do
+          it 'does not prune nil values' do
+            expect(codependent.new(a: 'hi', b: nil).to_hash).to eq(a: 'hi', b: nil, c: 'c')
+            expect(codependent.new(a: 'hi', b: nil, c: nil).to_hash).to eq(a: 'hi', b: nil, c: 'c')
+            expect(codependent.new(a: 'hi', b: nil).merge(c: nil).to_hash).to eq(a: 'hi', b: nil, c: nil)
+          end
+        end
+
+        describe '**' do
+          # Note: This test is an implementation detail of MRI and may not hold for
+          # other Ruby interpreters. But it's important to note in the test suite
+          # because it can be surprising for people unfamiliar with the semantics of
+          # double-splatting.
+          #
+          # For more information, see [this link][1]:
+          #
+          # [1]: https://github.com/hashie/hashie/issues/353#issuecomment-363294886
+          it 'prunes nil values because they are not set in the dash' do
+            dash = codependent.new(a: 'hi', b: nil)
+
+            expect(**dash).to eq(a: 'hi', c: 'c')
+          end
+        end
+      end
     end
   end
 end
@@ -482,6 +517,43 @@ class KeepingMash < Hashie::Mash
     expect(@bottom.new).to have_key(:echo)
     expect(@bottom.new).to_not have_key('echo')
   end
+
+  context 'exporting nil values' do
+    let(:test) do
+      Class.new(Hashie::Dash) do
+        property :foo
+        property :bar
+      end
+    end
+
+    describe '#to_h' do
+      it 'does not prune nil values' do
+        expect(test.new(foo: 'hi', bar: nil).to_h).to eq(foo: 'hi', bar: nil)
+      end
+    end
+
+    describe '#to_hash' do
+      it 'does not prune nil values' do
+        expect(test.new(foo: 'hi', bar: nil).to_hash).to eq(foo: 'hi', bar: nil)
+      end
+    end
+
+    describe '**' do
+      # Note: This test is an implementation detail of MRI and may not hold for
+      # other Ruby interpreters. But it's important to note in the test suite
+      # because it can be surprising for people unfamiliar with the semantics of
+      # double-splatting.
+      #
+      # For more information, see [this link][1]:
+      #
+      # [1]: https://github.com/hashie/hashie/issues/353#issuecomment-363294886
+      it 'prunes nil values because they are not set in the dash' do
+        dash = test.new(foo: 'hi', bar: nil)
+
+        expect(**dash).to eq(foo: 'hi')
+      end
+    end
+  end
 end
 
 describe SubclassedTest do